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It means "character" to differentiate between the role they play in SMP and the actual content creators.
c!Tubbo has nukes

I hate c!Dream so much he's literally the worst
c! by Peach<3 December 18, 2021
Related Words
C.O.W. c c hair C-row C.T. C.S. C.R. c notes C.O.T. c-town
The saddest man in the whole entire world.
the burden of the whole entire world is on my shoulders :C
:C by Fantastic Johnny C April 26, 2005
The majority of one's transcript should they opt into their local college of engineering.
I was certain I failed that Calculus III course, but thanks to the powers that be, I was blessed with a C-.
C- by Prometheus McSpanky April 18, 2014
Anime/cute section of 4chan. Calm and sporting a slow post rate, /c/ is a novelty compared to the other boards. Mainly an imageboard meant for posting images of anime/videogame girls that are cute/moe in nature.
"Wow, /c/ is on its way to its one millionth post! Sure took it long enough..."

"Gee, Bill! How'd you get so many pictures of Konata Izumi?"
"I found them on /c/."
/c/ by Iron Ladle May 26, 2009
C is for cookie, that's good enough for me.
when you take a bite from a cookie, it looks like a C
C by the real cookie monster July 27, 2005
A: I'm happy to see you again
Z: Me too c:
c: by Fainspirit November 5, 2018