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four on the floor 

refers to two things, in my experience.

1. a rhythm pattern where the percussive bass takes place on every downbeat. Common in house, trance, other forms of techno, disco, etc.

it is the opposite of a breakbeat, where the percussive bass can be anywhere, but is normally not on beats 2 and 4, and rarely on 3.

2. refers to a 4 speed transmittion with the shifter on the floor (as opposed to the dash or the column)
1. I dig that four-on-the-floor beat; it goes well with the bassline.

2. John wrecked the four-on-the-floor transmittion in his car by going 80 MPH and then quickly shifting into reverse.
four on the floor by karl December 26, 2004
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four on the floor 

n. Indicates that a car has a four-speed transmission, where the shifter is "on the floor" rather than on the steering column.
That old camaro had a 454 with four on the floor.
four on the floor by fizzle April 13, 2004

four on the floor 

a sex act in which one of the participants is on the floor with both arms and legs supporting them.
It was hot as hell...I was standing above her naked body and I had her from behind and she had four on the floor.

Four on the floor 

Sex game where either 4guys or 4women get naked and lie on the floor next to each other
They then have a jack off or jill off contest to see who cums first. The winner gets sucked or eaten by the other 3 cuntestants. The winner then sits on the side lines and the process repeats itself until they are down to the final 2 the.winner of this match up then gets sucked or eaten by the other 3 then he or she chooses who they want to fuck while the others watch while jacking or jilling off and cumming all over the intertwined couple.
Kiera went to a party last night where they played. Four on the floor. She said it was fun but she was exhausted when it was all over and she had to shower. 3 Jotimes to get the 'girl quim'out of her hair!

Four-On-The-Floor Beat 

A Simple Beat Made For Dancing. Found A Lot In Techno Pop Music.
This Song Has House Roots And A Four-On-The-Floor Beat I Can Chill And Dance To.
Four-On-The-Floor Beat by J.Babi January 28, 2011