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3.
A group devoted to the protection of the rights of E-4s and below in the Military to sham, blame problems on "I didn't know " or their leadership. The E-4 Mafia was created to bond lower enlisted against the tranny that is rank thru tenure and not based on the ability to lead.
Dude the Don call for a meeting of the E-4 Mafia we got to get that new 1SGT fired
by SPC IZZY January 05, 2010
 
1.
Upper-Junior Enlisted Army soldiers with loyalties in protecting all E-1 through E-4 against the seemingly unstoppable power of the NCO support channel.

Often the biggest contributor in spreading false information around the unit, and the leading reason Specialists evade doing mundane dirty work.
PV2: So, this is the fourth time in a row I got called out for K-P duty... I swear, I'm going to tell the Platoon Sergeant.

PFC: Shut your mouth, if any of the E-4 Mafia hear you, you're ass will be toasted. No sergeant could help you then.
by E-3 June 12, 2011
 
2.
The E-4 Mafia is the unofficial "gang" of the Specialist rank in the U.S. Army. Notorious for knowing more than the E-5 Sergeants and normally too much for their own good. Squad leaders are usually very pissed off when "secret" plans reach the E-4 Mafia before THEY even hear about it. Usually their intelligence comes from untrustworthy sources, but sometimes they get lucky.
GOD DAMN THAT E4-MAFIA, HOW MANY TIMES DO I HAVE TO TELL YOU NOT TO LISTEN TO THEM, PRIVATE!!
by Mikeymoped December 31, 2010