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Similar to the phrase "the bee's knees," but more crude. Implies that something is really cool, or sometimes used in sarcasm to imply that something is really not cool.
1 - Implying something is cool

- "That peach harvest sure was the bee's knees!"

- "Dude, that weed was the cockroach's testicles!"

2 - Sarcasm implying that something isn't really that cool

- "Hey man, check out this 1990 Ford Taurus I got used!" "Wow. That's just the cockroach's testicles isn't it...."
by C-Witt July 02, 2009