Slang term for 'testicles' - the 'under' part of 'undertones' refers to the position of the testicles in relation to the rest of the male reproductive system, plural because there are two testes.
tycho: Put on a pair of pants, Gabe!
gabe: Why, what's wrong with these short-shorts?
tycho: I can see your dystopian undertones, fer chrissake!
- penny arcade
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