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Car with automatic transmission. Also the selector in such a car

A car can be a prundler even though it might not have the traditional automatic transmission with torque converter. Any DSG, CVT, Hybrid that has a fully automatic mode for changing gear is a prundler.

Any car (usually electric) with fixed-ratio transmission is also a prundler.

To not be a prundler a car would need at least two forward gear ratios that can only be selected manually.

Origin:
From PRNDL -> PRNDLer -> PRuNDLer
(referring to the car itself)
Would you believe it, that Peugeot 407 is a prundler, look inside!

So there I was trundling along in my prundler...

(refering to the car part)
The prundler in my car is broken and now its stuck in reverse
prundler by towel406 June 21, 2011
Related Words

Prunelle 

A sassy girl who tells you how it is. But she does have a sweet side and has many friends. She's loud at times and if you get her mad enough, she will hit you. Hard.
Guy 1: Dude i don't think Prunelle likes me.
Guy 2: Why? What did you do?
Guy 1: I kept staring at her boobs and she punched me in the neck.
Prunelle by sexkitten593 January 7, 2012
When someone spends too much time in a shower, bath or swimming pool and ends up with soft, wrinkled skin. This then leads to the person looking like a Prune, hence the term.
Person 1: Man, that was a well deserved bath!

Person 2: I can tell - you're Pruning all over!
Pruning by Grbel08 August 16, 2011

Friend Pruning 

The act of removing multiple friends on one's friend list on social networking sites such as facebook and myspace
Why does Susan have less friends than usual on facebook?
Probably just doing a routine friend pruning on the people she doesn't like.
Friend Pruning by Ben Fong March 3, 2010
1. When one is "getting soft."

2. When one's manliness is quickly replaced with girly emotions.
3. When a person figuratively loses their balls
Man, that nigga Steffan just bout wet his pants when Josh barely pushed him to the side. He prunin' yo. Can't hang with us no more.

He's so whipped that he went through all that trouble to buy her a 300 dolla promise ring when he aint even getting laid. He prunin'.
Prunin' by Jwats Alyssa Steven April 28, 2011

Pruning the Hedges

The act of trimming your overgrown pubic hair
Tim: I just pruned the hedges. Phew, I really needed that.

Rob: What do you mean, pruning the hedges?

Tim: Look! See? I even made an arrow with it!

Rob: Dude, put that away! Nobody wants to see that!