a response to pretty much anything. it can express anger ('you know what? i dont care. titties in yo face') or a salutation ('goodbye and good luck. i wish you many titties in and around your face'). but mostly, it just describes one of the most awesome things that can happen to a guy- having titties in your face.
'dude, i hooked up with that chick i was hitting on last night.'
'titties in yo face?'
'yep, titties in my face.'
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